Update on the Brassicaceae Species Checklist
最新版本 由 Biodiversity Data Journal 發佈於 2021年8月31日 Biodiversity Data Journal

Here we present a revised species checklist for the Brassicaceae updated from Warwick SI, Francis, A., Al-Shehbaz IA (2006). Brassicaceae: Species checklist and database on CD-ROM. Plant Systematics and Evolution 259: 249─25. This update of the checklist was initiated on the basis of recent taxonomic and molecular studies on the Brassicaceae that have resulted in new species names, combinations and associated synonyms. New data has been added indicating tribal affiliations within the family and where type specimens have been designated. In addition, information from many early publications was checked and added to the database. The database now includes information on 14,980 taxa, 4,634 of which are currently accepted and divided into 340 genera and 52 tribes. A selected bibliography of recent publications on the Brassicaceae is included.

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Francis A, Lujan-Toro B E, Warwick S, Macklin J A, Martin S L (2021): Update on the Brassicaceae Species Checklist. v1.8. Biodiversity Data Journal. Dataset/Checklist. http://ipt.pensoft.net/resource?r=aafc-brassicaceae-checklist&v=1.8



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Checklist; Brassicaceae; mustard family; crucifer; global checklist



Ardath Francis
Agriculture and Agri-Food Canada
Beatriz E. Lujan-Toro
Biologist in Computational Biology
Agriculture and Agri-Food Canada
Suzanne Warwick
Retired scientist
Agriculture and Agri-Food Canada
James A. Macklin
Research Scientist
Agriculture and Agri-Food Canada
Sara L. Martin
Research Scientist
Agriculture and Agri-Food Canada


Beatriz E.. Lujan-Toro
Biologist in Computational Biology
Agriculture and Agri-Food Canada


Beatriz E. Lujan-Toro
Biologist in Computational Biology
Agriculture and Agri-Food Canada


Beatriz E. Lujan-Toro
Biologist in Computational Biology
Agriculture and Agri-Food Canada
Ardath Francis
Agriculture and Agri-Food Canada
Suzanne Warwick
Retired Scientist
Agriculture and Agri-Food Canada
James A. Macklin
Research Scientist
Agriculture and Agri-Food Canada
Sara L. Martin
Research Scientist
Agriculture and Agri-Food Canada

The checklist covers species, subspecies and varieties from the Brassicaceae family at a global scale.

界定座標範圍 緯度南界 經度西界 [-90, -180], 緯度北界 經度東界 [90, 180]

The database includes information on 14,980 taxa, 4,634 of which are currently accepted and divided into 340 genera and 52 tribes.

Kingdom  Plantae (plants)
Class  Equisetopsida (vascular plants)
Family  Brassicaceae (mustard or crucifer family)
起始日期 / 結束日期 1970-01-01 / 2019-01-01
  1. Al-Shehbaz IA (2006) The genus Sisymbrium in South America, with synopses of the genera Chilocardamum, Mostacillastrum, Neuontobotrys, and Polypsecadium (Brassicaceae). Darwiniana 44: 341-358.
  2. Al-Shehbaz IA (2007) Generic Limits of Dryopetalon, Rollinsia, Sibara, and Thelypodiopsis (Brassicaceae), and a Synopsis of Dryopetalon. Novon 17: 397-402. https://doi.org/10.3417/1055-3177(2007)17[397:glodrs]2.0.co;2
  3. Al-Shehbaz IA (2009) Two New North American Species of Draba (Brassicaceae): D. Heilii from New Mexico and D. Mulliganii from Alaska. Harvard Papers in Botany 14: 83-86. https://doi.org/10.3100/025.014.0112
  4. Al-Shehbaz IA (2010a) A Synopsis of the Genus Sibara (Brassicaceae). Harvard Papers in Botany 15: 139-147. https://doi.org/10.3100/025.015.0107
  5. Al-Shehbaz IA (2010b) A synopsis of the South American Lepidium (Brassicaceae). Darwiniana 48(2): 141-167. https://doi.org/10.3100/025.017.0202
  6. Al-Shehbaz IA (2012a) A generic and tribal synopsis of the Brassicaceae (Cruciferae). Taxon 61: 931-954. https://doi.org/10.1002/tax.615002
  7. Al-Shehbaz IA (2012b) Notes on Miscellaneous Species of the Tribe Thelypodieae (Brassicaceae). Harvard Papers in Botany 17: 3-10. https://doi.org/10.3100/025.017.0102
  8. Al-Shehbaz IA (2012c) A Synopsis of the South American Descurainia and Description of the Colombian New Species D. cleefii (Brassicaceae). Harvard Papers in Botany 17: 221-229. https://doi.org/10.3100/025.017.0202
  9. Al-Shehbaz IA (2013a) Clypeola is United with Alyssum (Brassicaceae). Harvard Papers in Botany 18: 125-128. https://doi.org/10.3100/025.018.0204
  10. Al-Shehbaz IA (2013b) A Revision of the Central American Genus Romanschulzia (Brassicaceae). Harvard Papers in Botany 18: 1-12. https://doi.org/10.3100/025.018.0102
  11. Al-Shehbaz IA (2014) A Synopsis of the Genus Noccaea (coluteocarpeae, Brassicaceae). Harvard Papers in Botany 19: 25-51. https://doi.org/10.1002/tax.615002
  12. Al-Shehbaz IA (2015) Typification of Himalayan Taxa of Brassicaceae (Cruciferae). Harvard Papers in Botany 20: 5-28. https://doi.org/10.3100/hpib.v20iss1.2015.n2
  13. Al-Shehbaz IA (2017a) Bengt-jonsellia (Brassicaceae), a New Genus from Madagascar. Harvard Papers in Botany 22: 1-3. https://doi.org/10.3100/hpib.v22iss1.2017.n1
  14. Al-Shehbaz, IA (2017b) Englerocharis cuzcoensis (Brassicaceae), a new species from Peru and a synopsis of Englerocharis. Novon 25: 399-402. https://doi.org/10.3417/d-17-00004
  15. Al-Shehbaz IA (2017c) Nomenclatural Adjustments in North American Arabis, Chinese Hilliella , and South American Neuontobotrys (Brassicaceae). Harvard Papers in Botany 22: 141-144. https://doi.org/10.3100/hpib.v22iss2.2017.n9
  16. Al-Shehbaz IA, German DA (2013) A synopsis of the genus Parrya (Brassicaceae). Kew Bulletin 68: 457-475. https://doi.org/10.1007/s12225-013-9450-4
  17. Al-Shehbaz IA, German DA (2014) A Synopsis of the Genus Braya (Brassicaceae). Harvard Papers in Botany 19: 161-174.
  18. Al-Shehbaz IA, German DA (2016) Three New Genera in the Tribe Euclidieae (Brassicaceae). Novon 25: 12-17. https://doi.org/10.3417/2016015
  19. Al-Shehbaz IA, Mayer MS (2013) New or Noteworthy Streptanthus (Brassicaceae) for the Flora of North America. Novon 18: 279-282. https://doi.org/10.3417/2007163
  20. Al-Shehbaz IA, Mulligan GA (2013) New or Noteworthy Species of Draba (Brassicaceae) from Canada and Alaska. Harvard Papers in Botany 18: 101-124. https://doi.org/10.3100/025.018.0203
  21. Al-Shehbaz I.A., Mummenhoff K. (2011). Stubendorffia and Winklera belong to the expanded Lepidium (Brassicaceae). Edinburgh Journal of Botany 68: 165–171. https://doi.org/10.1017/s0960428611000060
  22. Al-Shehbaz IA, Warwick SI (2007) Two New Tribes (Dontostemoneae and Malcolmieae) in the Brassicaceae (Cruciferae). Harvard Papers in Botany 12: 429-433. https://doi.org/10.3100/1043-4534(2007)12[429:tntdam]2.0.co;2
  23. Al-Shehbaz IA, Windham MD (2009). New and noteworthy North American Draba (Brassicaceae). Harvard Papers in Botany 12: 409–419. https://doi.org/10.3100/1043-4534(2007)12[409:nonnad]2.0.co;2
  24. Al-Shehbaz IA, Beilstein MA, Kellogg EA (2006) Systematics and phylogeny of the Brassicaceae (Cruciferae): An overview. Plant Systematics and Evolution 259: 89-120. https://doi.org/10.1007/s00606-006-0415-z
  25. Al-Shehbaz IA, Muñoz-Schick M, Morales V (2011b) The Present Status of Brassicaceae Taxa Described by Rodulfo and Federico Philippi. Harvard Papers in Botany 16: 279-291. https://doi.org/10.3100/
  26. Al-Shehbaz IA, Mutlu B, Dönmez AA (2007) The Brassicaceae (Cruciferae) of Turkey, Updated. Turkish Journal of Botany 31: 327-336.
  27. Al-Shehbaz IA, Cano A, Manchego MAC, Salariato DL (2018) Remarkable discoveries in the long-neglected and peruvian-endemic genus Machaerophorus (Brassicaceae). Phytotaxa 360: 114-124. https://doi.org/10.11646/phytotaxa.360.2.3
  28. Al-Shehbaz IA, Cano A, Trinidad H, Navarro E (2013) New species of Brayopsis, Descurainia, Draba, Neuontobotrys and Weberbauera (Brassicaceae) from Peru. Kew Bulletin 68: 219-231. https://doi.org/10.1007/s12225-013-9447-z
  29. Al-Shehbaz IA, German DA, Mummenhoff K, Moazzeni H (2014) Systematics, Tribal Placements, and Synopses of the Malcolmia S.L. Segregates (Brassicaceae). Harvard Papers in Botany 19: 53-71. https://doi.org/10.3100/hpib.v19iss1.2014.n4
  30. Al-Shehbaz IA, German D, Karl R, Jordon-Thaden I, Koch M (2011a) Nomenclatural adjustments in the tribe Arabideae (Brassicaceae). Plant Diversity and Evolution 129: 71-76. https://doi.org/10.1127/1869-6155/2011/0129-0044
  31. Alexander PJ, Windham MD, Govindarajulu R, Al-Shehbaz IA, Bailey CD (2010) Molecular Phylogenetics and Taxonomy of the Genus Thysanocarpus (Brassicaceae). Systematic Botany 35: 559-577. https://doi.org/10.1600/036364410792495926
  32. Alexander PJ, Windham MD, Beck JB, Al-Shehbaz IA, Allphin L, Bailey CD (2013) Molecular Phylogenetics and Taxonomy of the Genus Boechera and Related Genera (Brassicaceae: Boechereae). Systematic Botany 38: 192-209. https://doi.org/10.1600/036364413x661917
  33. ailey CD, Al-Shehbaz IA, Rajanikanth G (2007) Generic Limits in Tribe Halimolobeae and Description of the New Genus Exhalimolobos (Brassicaceae). Systematic Botany 32: 140-156. https://doi.org/10.1600/036364407780360166
  34. Cafferty S, Jarvis CE (2002) Typification of Linnaean plant names in Brassicaceae (Cruciferae). Taxon 51: 529-537. https://doi.org/10.2307/1554869
  35. Cecchi L (2011) A reappraisal of Phyllolepidium (Brassicaceae), a neglected genus of the European flora, and its relationships in tribe Alysseae. Plant Biosystems 145: 818-131. https://doi.org/10.1080/11263504.2011.580789
  36. Çetin Ö, Duran A, Martin E, Tuştaş S (2011) A taxonomic study of the genus Fibigia Medik. (Brassicaceae). African Journal of Biotechnology 11: 109-119. https://doi.org/10.5897/ajb11.302
  37. Chen H, Deng T, Yue J, Al-Shehbaz IA, Sun H (2016) Molecular phylogeny reveals the non-monophyly of tribe Yinshanieae (Brassicaceae) and description of a new tribe, Hillielleae. Plant Diversity 38: 171-182. https://doi.org/10.1016/j.pld.2016.04.004
  38. de Lange P, Heenan P, Houliston G, Rolfe J, Mitchell A (2013) New Lepidium (Brassicaceae) from New Zealand. PhytoKeys 24: 1-147. https://doi.org/10.3897/phytokeys.24.4375
  39. Esmailbegi S, Lysak MA, Rahiminejad MR, Mirtadzadini M, Mummenhoff K, Al-Shehbaz IA (2017a) A taxonomic revision of the genus Graellsia (Brassicaceae, tribe thlaspideae). Phytotaxa 313: 105-116. https://doi.org/10.11646/phytotaxa.313.1.7
  40. Esmailbegi S, Al-Shehbaz IA, Pouch M, Mandáková T, Mummenhoff K, Rahiminejad MR, Mirtadzadini M, Lysak MA (2018) Phylogeny and systematics of the tribe thlaspideae (Brassicaceae) and the recognition of two new genera. Taxon 67: 324-340. https://doi.org/10.12705/672.4
  41. Esmailbegi S, Lysak MA, Rahiminejad MR, Mirtadzadini M, Mummenhoff K, Al-Shehbaz I.A. (2017b). A taxonomic revision of the genus Pseudocamelina (Brassicaceae, tribe Thlaspideae). Phytotaxa 313: 117–129. https://doi.org/10.11646/phytotaxa.313.1.8
  42. German DA (2010) A checklist and system of the Cruciferae of Altai. Komarovia 6: 83-92.
  43. German DA (2012) Typification of some names in the Cruciferae, mostly from North Asia. Turczaninowia 15: 9-16.
  44. German DA (2013) Proposal to conserve the name Alyssum canescens (Ptilotrichum canescens, Stevenia canescens) (Cruciferae) with a conserved type. Taxon 62: 836-837. https://doi.org/10.12705/624.18
  45. German DA (2014a) New synonyms and combinations in Eurasian Brassicaceae (Cruciferae). Phytotaxa 173: 31-40. https://doi.org/10.11646/phytotaxa.173.1.2
  46. German DA (2014b) Some new and revised typifications in North Eurasian Cruciferae. Turczaninowia, 17: 29-41. https://doi.org/10.14258/turczaninowia.17.4.6
  47. German DA (2015) Cruciferae (Brassicaceae): Alternative treatment for the “Conspectus of the vascular plants of Mongolia” (2014). Turczaninowia 18: 39-67. https://doi.org/10.14258/turczaninowia.18.2.4
  48. German DA (2016) Taxonomic notes on miscellaneous Cruciferae. Turczaninowia 19: 129-135. https://doi.org/10.14258/turczaninowia.19.4.17
  49. German DA, Al-Shehbaz IA (2008a) Five Additional Tribes (Aphragmeae, Biscutelleae, Calepineae, Conringieae, and Erysimeae) in the Brassicaceae (Cruciferae). Harvard Papers in Botany 13: 165-170. https://doi.org/10.3100/1043-4534(2008)13[165:fatabc]2.0.co;2
  50. German DA, Al-Shehbaz IA (2008b) Dendroarabis, A New Asian Genus of Brassicaceae. Harvard Papers in Botany 13: 289-291. https://doi.org/10.3100/1043-4534-13.2.289
  51. German DA, Al-Shehbaz IA (2010) Nomenclatural novelties in miscellaneous Asian Brassicaceae (Cruciferae). Nordic Journal of Botany 28: 646-651. https://doi.org/10.1111/j.1756-1051.2010.00983.x
  52. German DA, Al-Shehbaz IA (2015a) Proposal to conserve the name Sisymbrium pumilum Stephan, non Lam. (Calymmatium pumilum, Olimarabidopsis pumila, Thellungiella pumila) (Cruciferae). Taxon 64: 849-850. https://doi.org/10.12705/644.21
  53. German DA, Al-Shehbaz IA (2015b) Typification of miscellaneous Brassicaceae (Cruciferae) from central and Middle Asia. Phytotaxa 221: 57-65. https://doi.org/10.11646/phytotaxa.221.1.5
  54. German DA, Al-Shehbaz IA (2018) A reconsideration of Pseudofortuynia and Tchihatchewia as synonyms of Sisymbrium and Hesperis, respectively (Brassicaceae). Phytotaxa 334: 95-98. https://doi.org/10.11646/phytotaxa.334.1.17
  55. German DA, Friesen NW (2014) Shehbazia (Shehbazieae, Cruciferae), a new monotypic genus and tribe of hybrid origin from Tibet. Turczaninowia 17: 17-23. https://doi.org/10.14258/turczaninowia.17.4.3
  56. German DA, Tekin M, Španiel S, Marhold K, Alshehbaz IA (2016) A brief taxonomic revision of Physoptychis (Alysseae, Brassicaceae). Phytotaxa 258: 75-82. https://doi.org/10.11646/phytotaxa.258.1.5
  57. Hall JC, Tisdale TE, Donohue K, Wheeler A, Al-Yahya MA, Kramer EM (2011) Convergent evolution of a complex fruit structure in the tribe Brassiceae (Brassicaceae). American Journal of Botany 98: 1989-2003. https://doi.org/10.3732/ajb.1100203
  58. Hao G, Al-Shehbaz IA, Ahani H, Liang Q, Mao K, Wang Q, Liu J (2017) An integrative study of evolutionary diversification of Eutrema (Eutremeae, Brassicaceae). Botanical Journal of the Linnean Society 184: 204-223. https://doi.org/10.1093/botlinnean/box024
  59. Heenan PB (2017) A taxonomic revision of Cardamine L. (Brassicaceae) in New Zealand. Phytotaxa 330: 1-154. https://doi.org/10.11646/phytotaxa.330.1.1
  60. Heenan PB, Goeke DF, Houliston GJ, Lysak MA (2012) Phylogenetic analyses of ITS and rbcL DNA sequences for sixteen genera of Australian and New Zealand Brassicaceae result in the expansion of the tribe Microlepidieae. Taxon 61: 970-979. https://doi.org/10.1002/tax.615004
  61. Hildebrand TJ, Al-Shehbaz IA (2017) Terraria haydenii (Thelypodieae, brassicaceae), a new mustard genus and species from the west desert region of North America’s Great Basin. Phytotaxa 323: 128-142. https://doi.org/10.11646/phytotaxa.323.2.2
  62. Hu H, Al-Shehbaz IA, Sun Y, Hao G, Wang Q, Liu J (2015) Species delimitation in Orychophragmus (Brassicaceae) based on chloroplast and nuclear DNA barcodes. Taxon 64: 714-726. https://doi.org/10.12705/644.4
  63. Jordon-Thaden I, Hase I, Al-Shehbaz IA, Koch MA (2010) Molecular phylogeny and systematics of the genus Draba (Brassicaceae) and identification of its most closely related genera. Molecular Phylogenetics and Evolution 55: 524-540. https://doi.org/10.1016/j.ympev.2010.02.012
  64. Koch MA (2012) Mid-miocene divergence of Ionopsidium and Cochlearia and its impact on the systematics and biogeography of the tribe Cochlearieae (Brassicaceae). Taxon 61: 76-92. https://doi.org/10.1002/tax.611006
  65. Koch, MA, Al-Shehbaz, IA (2009) Molecular systematics and evolution: Recognition of infrafamiliar taxa: the tribal system. In: Gupta, SK (Ed) Biology and breeding of crucifers. CRC Press, Boca Raton, 1-18. https://doi.org/10.1201/9781420086096
  66. Koch MA, Karl R, German DA, Al-Shehbaz IA (2012) Systematics, taxonomy and biogeography of three new Asian genera of Brassicaceae tribe Arabideae: An ancient distribution circle around the Asian high mountains. Taxon 61: 955-969. https://doi.org/10.1002/tax.615003
  67. Mandáková T, Mummenhoff K, Al-Shehbaz IA, Mucina L, Mühlhausen A, Lysak MA (2012) Whole-genome triplication and species radiation in the southern African tribe Heliophileae (Brassicaceae). Taxon 61: 989-1000. https://doi.org/10.1002/tax.615006
  68. Marhold K (2011) Brassicaceae. In: Euro+Med Plantbase - the information resource for Euro-Mediterranean plant diversity. http://ww2.bgbm.org/EuroPlusMed/ [accessed 14.11.2016]
  69. Marhold K, Kempa M, Al-Shehbaz IA (2015) Lectotypification of names of Himalayan Brassicaceae taxa currently placed in the genus Cardamine. PhytoKeys 50: 9-23. https://doi.org/10.3897/phytokeys.50.5080
  70. Marhold K, Lihová J, Al-Shehbaz IA, Kudoh H (2007) The correct interpretation and lectotypification of the name Cardamine fallax (Brassicaceae). Journal of Plant Research 120: 655-660. https://doi.org/10.1007/s10265-007-0107-y
  71. Marhold K, Ŝlenker M, Kudoh H, Zozomová-Lihová J (2016) Cardamine occulta, the correct species name for invasive Asian plants previously classified as C. flexuosa, and its occurrence in Europe. PhytoKeys 62: 57-72. https://doi.org/10.3897/phytokeys.62.7865
  72. Moazzeni H, Al-Shehbaz IA, German DA, Assadi M, Müller J, Joharchi MR, Memariani F (2018) A taxonomic revision of the genus Aethionema S.L. (Brassicaceae) in Iran. Phytotaxa 356: 241-266. https://doi.org/10.11646/phytotaxa.356.4.1
  73. Nikolov LA, Shushkov P, Nevado B, Gan X, Al-Shehbaz IA, Filatov D, Bailey CD, Tsiantis M (2019) Resolving the backbone of the Brassicaceae phylogeny for investigating trait diversity. New Phytologist 222: 1638-1651. https://doi.org/10.1111/nph.15732
  74. Özüdoğru B, Al-Shehbaz IA, Mummenhoff K (2017) Tribal assignment of Heldreichia Boiss. (Brassicaceae): evidence from nuclear ITS and plastidic ndhF markers. Plant Systematics and Evolution 303: 329-335. https://doi.org/10.1007/s00606-016-1374-7
  75. Parolly G, Nordt B, Bleeker W, Mummenhoff K (2010) Heldreichia Boiss. (Brassicaceae) revisited: A morphological and molecular study. Taxon 59: 187-202. https://doi.org/10.1002/tax.591018
  76. Prina AO (2009) Taxonomic review of the genus Crambe sect. Crambe (Brassicaceae, Brassiceae). Anales del Jardín Botánico de Madrid 66: 7-24. https://doi.org/10.3989/ajbm.2186
  77. Prina AO, Martínez-Laborde JB (2008) A taxonomic revision of Crambe section Dendrocrambe (Brassicaceae). Botanical Journal of the Linnean Society 156: 291-304. https://doi.org/10.1111/j.1095-8339.2007.00708.x
  78. Ranjbar M, Karami S, Rostami M (2008) A revision of Fibigia sect. Purpureae (Brassicaceae, Alysseae) in Iran, and the description of three new species. Biological Diversity and Conservation 7: 32-43. https://doi.org/10.11646/phytotaxa.258.1.5
  79. Salariato DL, Al-Shehbaz IA (2014) Zuloagocardamum (Brassicaceae: Thelypodieae) a New Genus from the Andes Highlands of Northern Argentina. Systematic Botany 39: 563-577. https://doi.org/10.1600/036364414x680898
  80. Salariato DL, Zuloaga FO (2015) Taxonomic placement of Onuris hauthalii (Brassicaceae: Eudemeae), based on morphology and multilocus species tree analyses, and the recognition of the new genus Alshehbazia. Kew Bulletin 70: 1-8. https://doi.org/10.1007/s12225-015-9602-9
  81. Salariato DL, Al-Shehbaz IA, Zuloaga FO (2018) Reinstatement of the Southern Andean Genus Stenodraba (Brassicaceae) Based on Molecular Data and Insights from its Environmental and Geographic Distribution. Systematic Botany 43: 35-52. https://doi.org/10.1600/036364418x697085
  82. Salariato DL, Zuloaga FO, Al-Shehbaz IA (2013a) Molecular phylogeny of Menonvillea and recognition of the new genus Aimara (Brassicaceae: Cremolobeae). Taxon 62: 1220-1234. https://doi.org/10.12705/626.6
  83. Salariato DL, Zuloaga FO, Al-Shehbaz IA (2013b) Revision and tribal placement of the Argentinean genus Parodiodoxa (Brassicaceae). Plant Systematics and Evolution 299: 305-316. https://doi.org/10.1007/s00606-012-0722-5
  84. Salariato DL, Zuloaga FO, Al-Shehbaz IA (2014) A revision of the genus Menonvillea (Cremolobeae, Brassicaceae). Phytotaxa 162: 241-292. https://doi.org/10.11646/phytotaxa.162.5.1
  85. Salariato DL, Zuloaga FO, Al-Shehbaz IA (2015) A taxonomic revision of the genus Xerodraba (Eudemeae, Brassicaceae). Phytotaxa 207: 39-67. https://doi.org/10.11646/phytotaxa.207.1.2
  86. Salariato DL, Zuloaga FO, Cano A, Al-Shehbaz IA (2015) Molecular phylogenetics of tribe Eudemeae (Brassicaceae) and implications for its morphology and distribution. Molecular Phylogenetics and Evolution 82: 43-59. https://doi.org/10.1016/j.ympev.2014.09.030
  87. Salariato DL, Cueva Manchego MA, Cano A, Al-Shehbaz IA (2019) Phylogenetic placement of the Peruvian-endemic genus Machaerophorus (Brassicaceae) based on molecular data and implication for its systematics. Plant Systematics and Evolution 305: 77-87. https://doi.org/10.1007/s00606-018-1553-9
  88. Salariato DL, Zuloaga FO, Franzke A, Mummenhoff K, Al-Shehbaz IA (2016) Diversification patterns in the CES clade (Brassicaceae tribes Cremolobeae, Eudemeae, Schizopetaleae) in Andean South America. Botanical Journal of the Linnean Society 181: 543-566. https://doi.org/10.1111/boj.12430
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替代的識別碼 94308742-058c-46d5-b763-06e9207a6b15